生长素在烟草表皮毛中的极性运输及对细胞伸长的影响
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湖南省教育厅重点科研项目(18A104)


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    摘要:

    利用生长素极性运输转运蛋白PIN1与增强绿色荧光蛋白EGFP的融合,对PIN1进行了荧光标记,并以烟草表皮毛为模式,开展了PIN1对生长素极性运输及对细胞伸长生长影响的研究。采用DNA重组技术,将融合标记基因EGFP–PIN1置于拟南芥表皮毛特异表达基因GL2的启动子调控下,构建成含GL2pro::EGFP–PIN1的Ti质粒,以根癌农杆菌叶盘共培转化法将重组标记基因转化至烟草WS38中,筛选鉴定出多株转基因烟草。通过对这些转基因烟草表皮毛进行显微荧光观察,结果发现,标记的绿色荧光信号集中分布在表皮毛细胞的间隔区,表现为明显的极性分布现象。用生长素极性运输抑制剂三碘苯甲酸(TIBA)处理后,表皮毛的伸长生长受到抑制,细胞中荧光的分布极性减弱。说明生长素在烟草表皮毛中的极性分布对烟草表皮毛伸长起关键作用,抑制生长素的极性运输不只抑制表皮毛的细胞伸长,同时还影响到生长素极性运输蛋白PIN的极性分布。

    Abstract:

    In this study, the enhanced green fluorescent protein(EGFP) was combined with auxin polar transporter PIN1 and made it fluorescently labeled. The effects of PIN1 on auxin polar transport and cell elongation growth were studied in tobacco trichome. The fusion label gene EGFP-PIN1 was recombined under the promoter of the trichome specific expression gene GL2 from Arabidopsis thaliana to enable it specific expression in the transgenic. The recombinant GL2pro::EGFP-PIN1 was constructed in the Ti plasmid pEGAD and then transformed into tobacco WS38 by Agrobacterium tumefaciens mediated leaf disc co-culture transformation. Several transgenic tobacco plantlets were screened out and identified. The fluorescence observation of these transgenic tobacco trichome showed that the labeled green fluorescence signals appeared in the interval area of the trichome cells with an obvious polarity distribution. It enables the polarity transport of auxin in trichrome cells and forms a polarity distribution of axuin. After treatment with the auxin polar transport inhibitor triiodobenzoic acid (TIBA), the elongation growth of trichome was significantly inhibited and the polarity distribution of fluorescence in cells was also weakened. Thus, we conclude that the polarity distribution of auxin in tobacco trichome plays a key role in the elongation of tobacco trichome. Inhibition of auxin polar transport not only inhibited cell elongation of trichome, but also affected the polarity distribution of auxin polar transport protein PIN.

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潘嘉立,赵燕,黄丽华,高晗,黄妤,陈金军,张学文.生长素在烟草表皮毛中的极性运输及对细胞伸长的影响[J].湖南农业大学学报:自然科学版,2020,46(6):.

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  • 在线发布日期: 2020-12-23
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